Emilia Fox as Sylvia Brown

Victoria Wood as Nana

Emma Watson as Pauline Fossil

Yasmin Paige as Petrova Fossil

Lucy Boynton as Posy Fossil

Marc Warren as Mr. Simpson

Richard Griffiths as Great Uncle Matthew

Lucy Cohu as Theo Dane

Gemma Jones as Dr. Jakes

Harriet Walter as Dr. Smith

Eileen Atkins as Madame Fidolia

Heather Nicol as Winifred Bagnall

Mary Stockley as Miss Jay

Teresa Churcher as Clara

Skye Bennett as Young Sylvia

Peter Bowles as Sir Donald Houghton

Don Gallagher as Mr. French

Emma Darwall Smith as Titania

Isabella Parriss as Cobweb

Adrian Lester as Mr. Sholsky

Tim Wallers as Mr. Montague

Gresby Nash as Leopold Neville

Ebe Sievwright as Valentin Manoff

Ruth Brill as Featured Dancer

Natasha Mould as Featured Dancer

Jack Thorpe-Baker as Featured Dancer

Jade Longley as Moth